The papers included in this volume were presented at the VIth International Ephemeroptera Conference and the Xth International Symposium on Plecoptera, and at a Joint Symposium on Applied Aspects in the study of Mayflies and Stoneflies. One additional paper has been included, a review of the studies of on mayflies.
Among the various groups of aquatic insects, mayflies (Ephemeroptera) are of special interest for professional limnologists and entomological researchers as well as for naturalists in general and even the dedicated fly angler. Identification has traditionally been considered difficult and implementation in environmental monitoring and freshwater management has led to an ever increasing demand for exact information on taxonomy and ecology. The present handbook is designed to provide for the first time an up-to-date standard work for Ephemeroptera identification, including last instar larvae (nymphs), subimago (dun), male and female imagines. Recent changes in nomenclature are discussed in detail as well as gaps in current knowledge and probable pitfalls concerning the reliable identification of all taxa known so far from the region. Keys are provided for genera and introductory chapters characterize every family and genus. Species accounts follow a common format providing a synonymy, characters for identification (including literature references), remarks (on type material, variation, confusing or extralimital species) and short information on biology and distribution pattern. Male genitalia are illustrated by micrographs and line drawings, REM photographs of the egg chorionic structure are provided for genera and selected species. Habitus of larvae and imagines are for most genera illustrated by colour photographs. The geographical area covered is Europe including the European part of Russia, the mediterranean islands and North Africa. Short additional information is provided for adjacent parts of the western Palaearctic Region. A comprehensive index, check-list and distribution catalogue (following the widely adopted concept of Illies’ Limnofauna Europaean) allow for quick information on all species recorded so far from Europe.
seem as appropriate now as the original balance was when Dr A. D. Imms' textbook was first published over fifty years ago. There are 35 new figures, all based on published illustrations, the sources of which are acknowledged in the captions. We are grateful to the authors concerned and also to Miss K. Priest of Messrs Chapman & Hall, who saved us from many errors and omissions, and to Mrs R. G. Davies for substantial help in preparing the bibliographies and checking references. London O. W. R. R. G. D. May 1976 Part III THEORDERSOFINSECTS THE CLASSIFICATION AND PHYLOGENY OF INSECTS The classification of insects has passed through many changes and with the growth of detailed knowledge an increasing number of orders has come to be recognized. Handlirsch (1908) and Wilson and Doner (1937) have reviewed the earlier attempts at classification, among which the schemes of Brauer (1885), Sharp (1899) and Borner (1904) did much to define the more distinctive recent orders. In 1908 Handlirsch published a more revolutionary system, incorporating recent and fossil forms, which gave the Collembola, Thysanura and Diplura the status of three independent Arthropodan classes and considered as separate orders such groups as the Sialoidea, Raphidioidea, Heteroptera and Homoptera. He also split up the old order Orthoptera, gave its components ordinal rank and regrouped them with some of the other orders into a subclass Orthopteroidea and another subclass Blattaeformia.
The idea of writing this book was conceived when, in the late 1960s, I began teaching a senior undergraduate class in general entomology. I soon realized that there was no suitable text for the class I intended to give. The so-called "general" or "introductory" texts reflected the traditional taxonomic approach to entomology and contained relatively little information on the physiology and ecology of insects. This does not mean that there were no books containing such information. There were several, but these were so specialized and de tailed that their use in an introductory class was limited. I hold a strong belief that an undergraduate general entomology course should provide a balanced treatment of the subject. Thus, although some time should be devoted to taxonomy, including identification (best done in the laboratory, using primar ily material which students themselves have collected, supplemented with specimens from the general collection), appropriate time should be given also to discussion of the evolution, development, physiology, and ecology of in sects. In the latter category I include the interactions between insects and Man because it is important to stress that these interactions follow normal ecological principles. Naturally, the format of this book reflects this belief. The book has been arranged in four sections, each of which necessarily overlaps with the others.
The book has general biological significance due to usage of the new non-ranking nomenclature and the rational layout of taxonomic text, which can be qualified as post-Linnaean systematics. While after the works by Lameere and Hennig, non-ranking classifications became widely used, this book represents the first experience of consistently non-ranking classification, including taxa of low taxonomic level (i.e. taxa traditionally regarded as genera, families et al.). In contrast to other recent attempts to elaborate a non-ranking nomenclature not contradictory to the International Code of Zoological Nomenclature, this one appears to be successful and can be applied to any zoological taxa (although its application in botany could be possible only after further elaboration). Biology is currently going through a crisis, which causes some investigators to use such non-scientific methods of reconstructing phylogeny as parsimony analysis. The author believes that the new method of phylogeny description and reconstruction used in this book will help indicate a way out of this crisis.
In his preface to early editions of this book, the late Dr. A. D. Imms said that he intended it to be an elementary account of entomology as a branch of general biology. He had especiaHy in mind the needs of university students of zoology and agriculture, as weH as those intending later to specialize in entomology, and he suggested that the book might also interest teachers of advanced biology in schools. These general aims and the balance between the different aspects of the subject have changed linIe in this and in our previous revision. We have, however, tried to bring the present edition up to date on the lines of our revised tenth edition of Imms' General Textbook 0/ Entomology, published in 1977. The text has been entirely re-set and eleven illustrations have been replaced by new figures. The same orders of insects are recognized as in the last edition, but the sequence in which the Endopterygote groups appear has been changed to reflect more accurately their probable evolutionary relationships. Many small changes and some addi tions have been made in the physiological sections, the chapter on the origin and phylogeny of insects has been rewritten, and a new bibliography provides a selection of modern references for the in tending specialist. It has been our object to make these alterations without materially increasing the length of the book or its level of difficulty.
Extensively revised, the Second Edition continues to offer senior undergraduate students a well-balanced treatment of all major areas in entomology. This edition features coverage of the new phylogenies for most of the insect orders